WSEAS Transactions on Mathematics
Print ISSN: 1109-2769, E-ISSN: 2224-2880
Volume 25, 2026
Nonlinear Dynamics and Bifurcation Structure of a Stage-Structured
Theta-Ricker Map with Cross-Stage Feedback
Authors: ,
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Abstract: This study surveyed the nonlinear dynamics of a discrete-time, stage-structured population model
merging demographic memory through adult survival. The model develops the classical θ–Ricker formulation
by introducing cross-stage feedback between juvenile and adult compartments governed by a set of biologically
interpretable parameters. The analytical results show that the system admits a unique positive equilibrium, the
local stability of which is determined by the trace and determinant of the Jacobian matrix. Loss of stability occurs
via two primary mechanisms: flip (period doubling) and Neimark–Sacker bifurcations. Explicit conditions for
both bifurcations are derived, and the first Lyapunov coefficient is computed to characterize the direction and
criticality of the Neimark–Sacker (NS) transition. Numerical experiments, including one-parameter bifurcation
diagrams and two-parameter Lyapunov maps, confirmed the analytical predictions. Increasing the reproductive
rate or cross-stage feedback leads to successive period doubling and chaos, whereas higher maturation efficiency
and adult survival expand the stability domain. The results revealed that demographic memory acts as an internal
damping mechanism that delays the onset of instability and suppresses chaotic oscillations. Overall, this study
provides a coherent analytical–numerical framework for understanding how feedback and memory interact to
shape complex population dynamics in stage-structured ecological systems.
Keywords:
Stage-structured population model, θ–Ricker map, cross-stage feedback, demographic memory, flip bifurcation, Neimark–Sacker bifurcation
Pages: 16-28
DOI: 10.37394/23206.2026.25.3